FRAGMENTA PALAEONTOLOGICA HUNGARICAVolume 35 Budapest, 2018 pp. 3–59

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Volume 35 Budapest, 2018 pp. 3–59

First supplement to the catalogue of

invertebrate and vertebrate palaeontological type specimens of the Hungarian Natural History Museum: 2008–2018

Alfréd Dulai¹, Mihály Gasparik¹, Zoltán Szentesi¹ & József Pálfy^{2, 3}

1Department of Palaeontology and Geology, Hungarian Natural History Museum, H-1083 Budapest, Ludovika tér 2, Hungary. E-mails: dulai.alfr ed@nhmus.hu,

gasparik.mihaly@nhmus.hu, szentesi.zoltan@nhmus.hu;

2MTA–MTM–ELTE Research Group for Palaeontology, H-1083 Budapest, Ludovika tér 2, Hungary.

E-mail: palfy.jozsef@nhmus.hu;

3Department of Geology, Eötvös Loránd University, H-1117 Budapest, Pázmány Péter sétány 1/C, Hungary.

E-mail: palfy@elte.hu

Summary – A comprehensive palaeontological type catalogue of the Hungarian Natural History Museum (Budapest) was published in 2008. In the following ten years several new type specimens were deposited in the collection, both invertebrates and vertebrates, from microscopic radiolarians to large dinosaur bone remains. Triassic radiolarians and ammonoids, Jurassic brachiopods, Cenozoic molluscs, and Cretaceous vertebrates represent the most dynamically growing parts of the type collection. Th is supplement to the original catalogue contains the data of 175 species and subspecies, of which 148 are new taxa for the type catalogue. Among them 125 are represented by holotypes, and an additional 199 inventory lots of their paratypes; 23 new taxa are represented only by paratypes, in an additional 74 inventory lots. Th e vast majority of these taxa were recently described (between 2008 and 2018), although a few of them are previously established taxa with type material that was newly recognized to exist in our collection. Th is supplement also contains the data of four recently designated neotypes, as well as 16 rediscovered old type specimens which were reported as missing in the 2008 catalogue. We also include the known revisions and illustration of 14 previously established species and subspecies (in three cases with lectotype designation).

At the end of 2018, the invertebrate and vertebrate palaeontological collection of the Hungarian Natural History Museum holds more than 2500 inventory lots of diff erent type specimens, including name-bearing types of 1056 nominal species and subspecies.

Key words – holotype, invertebrates, lectotype, neotype, new taxa, palaeontology, paratype, ver- tebrates

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Introduction 5

Catalogue of invertebrate type specimens 10

1. Triassic types 10

1.1. Triassic Radiolaria 10

1.2. Triassic Gastropoda 18

1.3. Triassic Cephalopoda 18

2. Jurassic types 21

2.1. Jurassic Bivalvia 21

2.2. Jurassic Gastropoda 22

2.3. Jurassic Cephalopoda 24

2.4. Jurassic Brachiopoda 25

3. Cretaceous types 26

3.1. Cretaceous Cephalopoda 26

3.2. Cretaceous Brachiopoda 26

3.3. Cretaceous Crinoidea 26

3.4. Cretaceous Echinoidea 27

4. Eocene types 27

4.1. Eocene Foraminiferida 27

4.2. Eocene Polyplacophora 28

4.3. Eocene Brachiopoda 28

5. Oligocene types 29

5.1. Oligocene Gastropoda 29

5.2. Oligocene Decapoda 29

6. Miocene types 32

6.1. Miocene Bivalvia 32

6.2. Miocene Gastropoda 32

6.3. Miocene Brachiopoda 34

6.4. Miocene Decapoda 34

6.5. Miocene Echinoidea 34

Catalogue of vertebrate type specimens 35

7. Amphibia 35

8. Reptilia 36

9. Aves 46

10. Mammalia 46

Index of species and subspecies names 49

References 52

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INTRODUCTION

Aft er moving of the palaeontological collections to a new location in 2004–

2006, and the development of a digital collection database, the fi rst catalogue of palaeontological type specimens of the Hungarian Natural History Museum (HNHM) was published by Pálfy et al. (2008). Some additional data on the types and the collections were summarized by Pálfy (2009). Th e history and development of the type catalogue was described in detail in Pálfy et al. (2008).

Th e catalogue provided a comprehensive listing of invertebrate and vertebrate type specimens published between 1841 and February 2008. Th e palaeobotani- cal type specimens are historically part of the collection of the Department of Botany at the HNHM (Hably & Szakály 1989; Hably et al. 2001).

Th e catalogue listed type specimens of 1181 fossil species-group taxa (986 invertebrates and 195 vertebrates). Discounting the specimens that are missing, lost or have been transferred to other institutions, the catalogue included prima- ry types of 919 taxa available in the collection of the HNHM. At that time 2017 existing inventory lots contained type specimens.

Th e present work supplements Pálfy et al.’s (2008) catalogue, including the type specimens published between February 2008 and December 2018, as well as those specimens which were regarded as missing types in the original catalogue, but later were rediscovered in the collection. We also include those previously published types which were revised or re-illustrated between 2008 and 2018.

Several type specimens were recently discovered in some historical collections or older materials, or were donated to the HNHM (e.g., Papp 1907; Kutassy 1933;

Somos & Kókay 1960; Steininger et al. 1973).

Following the structure of the original catalogue, this supplement also in-

cludes primary, name-bearing types (holotypes, syntypes, lectotypes, neotypes)

and secondary types (paratypes, paralectotypes). It is subdivided into separate

parts for invertebrate and vertebrate types. Th e basic information of taxa and

their type specimens is listed in slightly diff erent formats for invertebrates and

vertebrates. An individual catalogue entry typically corresponds to an invento-

ry lot that contains the type specimen(s) of a taxon, identifi ed by an inventory

number. If a taxon is represented by more than one type specimen, separate en-

tries are listed where diff erences exist in either the inventory number, locality,

or illustration of specimen. Types under more than one inventory number are

merged into a single entry if all data fi elds other than the inventory number are

identical.

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Th e format for invertebrate entries is as follows:

Genus species Author, year

[or Genus (Subgenus) species subspecies Author, year]

Status of type, inventory number

[Period and/or epoch], standard or regional stage, formation Locality, town, country (country is listed only if other than Hungary) Reference, page, text-fi gure, plate, fi gure (see code numbers in References) Revised name, reference to revising work

Remarks

Th e format for pre-Pliocene vertebrate entries is as follows:

Status of type, inventory number Part of fossil

Stage, epoch; formation Locality, town, country

Reference, page, text-fi gure, plate, fi gure Revised name, reference to revising work Remarks

Th e format for Pliocene and Quaternary vertebrate entries is as follows:

Status of type, inventory number Part of fossil

Epoch subdivided (i.e. Early/Middle/Late Pliocene or Pleistocene) Locality, town, country

Reference, page, text-fi gure, plate, fi gure Revised name, reference to revising work Remarks

Th e invertebrate catalogue entries are arranged primarily by geologic age, secondarily by major taxonomic groups, and then the original proposed bi- or trinomens are listed alphabetically within each subdivision.

Both the ages and the taxonomic groups that defi ne the subdivisions were selected for practical reasons rather than strictly using certain levels in their hier- archical structure. Th e following age subdivisions, ranging in rank from period to epoch, are used: Triassic, Jurassic, Cretaceous, Eocene, Oligocene, and Miocene.

Secondary, taxonomic subdivisions use the following, well-known groups, arbi- trarily taken from higher taxa at the phylum, class, or order level: Radiolaria, Foraminiferida, Polyplacophora, Bivalvia, Gastropoda, Cephalopoda, Brachio- poda, Decapoda, Crinoidea, Echinoidea.

Th e catalogue of vertebrate types is arranged by higher taxa, containing sub-

divisions for Amphibia, Reptilia, Aves, and Mammalia.

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Taxon names for the types are given as bi- or trinomens as they were intro- duced by the author in the original description. Th e original spelling is retained, even though current rules of ICZN (see Article 32) require avoiding the diacriti- cal marks of non-standard Latin characters (commonly occurring in taxon names derived from Hungarian person or place names). However, preserving the origi- nal spelling in the catalogue may help tracing a taxon in the literature. Necessary corrections and emendations of originally incorrect spelling of species names are refl ected in the index of species and subspecies names.

If types of a taxon require more than one entry, the holotype or lectotype is listed fi rst. For types of equal status (e.g. syntypes, paratypes, paralectotypes), the entries are arranged in ascending order of their inventory number. In merged entries with several specimens of identical attributes, the inventory numbers are given in a list separated by commas.

If possible, an age assignment for invertebrate specimens is given at the stage level. Standard stages are used (Gradstein et al. 2012) except for the Oligocene and Miocene, where the regional stages of the Central Paratethys time scale are applied (Piller et al. 2007). Although some stages are known to strad- dle standard epoch boundaries, for sake of a simplifi ed hierarchy of age units, these boundary problems are sidestepped. Th us the Egerian is included in the Oligocene, and the Pannonian is taken to form part of the Miocene. Original age assignments are revised if necessary, where up-to-date stratigraphic information is available from the type locality.

Th e ages of Pliocene and Pleistocene vertebrates are given using a three- fold (i.e. Early–Middle–Late) subdivision of the epochs, derived by simplify- ing the biochronological framework of Kretzoi (1969) and Jánossy (1986), with the necessary modifi cations to account for diff erent concepts in placing the Pliocene/Pleistocene boundary. Age assignment of pre-Pliocene vertebrates fol- lows the concepts applied to invertebrates.

Th e lithostratigraphic unit from which a type specimen was collected is given on the basis of Császár (1997) for Hungary; in other countries the rel- evant references were consulted for information on lithostratigraphy. Pliocene and younger vertebrates are not assigned to formations as they are most com- monly found in fi ssure fi lls and cave deposits, not forming conventional lithos- tratigraphic units.

In the locality fi eld, geographic names are given in their usual Hungarian form, for ease of fi nding them on local maps. Th e English translation of some common vernacular terms in geographic names (oft en hyphenated to form the latter part of a proper name) is listed as follows (Hungarian words in italics): árok – gully; domb – hill; hegy – mountain; kút – well; mező – meadow; puszta – farm;

völgy – valley. Geological terms for localities, such as outcrop (“feltárás”), quarry

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(“kőfejtő”), brickyard (“téglagyár”), coal mine (“szénbánya”) are given in English.

Name of town refers to the municipality to which the locality belongs, in some cases this is not the nearest settlement.

As the majority of the types were collected at localities within the present- day borders of Hungary (385 inventory lots), no country name is listed for them.

Some specimens are from sites in historical Hungary that are now in the terri- tory of Slovakia (7) or Romania (1). Th eir town is listed using the current offi cial name, with the Hungarian name given in parentheses. Approximately one-quar- ter of the types are from other countries, including Turkey (97), Austria (16), Russia (8), France (8), and Poland (3).

Reference is given to the original description, using a fi ve or six character code, found in the reference list at the end of each item. Th e reference code is composed of the fi rst three letters of the fi rst author’s name (or an otherwise meaningful three-letter abbreviation of the name), the last two digits of the year of publication, and, if needed, an extra letter to distinguish works from the same year. If the original description appeared in Hungarian as well as in a foreign language, both page numbers are given, the foreign one fi rst and the Hungarian one in parentheses.

Illustrations (text-fi gures and/or fi gures on plates) accompanying the origi- nal description are quoted at the appropriate entry, if the specimen is identifi ed as the fi gured one.

If we became aware of a published revision, either nomenclatural, or changing the species-level identifi cation or the generic assignment of the taxon concerned, we include the revised name and provide the reference code of the revising work.

Any additional information may appear as remarks in the last line of an item.

Remarks may include reference to a diff erent inventory number in the original pub- lication, notes on the type designation, or the date of rediscovery in the collection.

To facilitate fi nding information on taxa, the catalogue is followed by an index of all invertebrate and vertebrate species and subspecies names. Here an original name (using emended spelling if needed) is followed by the originally published bi- or trinomen, the author’s name, and the page number on which the fi rst catalogue entry of this taxon appears.

Although signifi cant care was taken to compile a comprehensive catalogue

that contains accurate information, errors and omissions are almost inevitable in

an endeavour of this scale. Th erefore users of this catalogue are encouraged to re-

port any suggested correction and/or addition to the curators of the Department

of Palaeontology and Geology.

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*

Acknowledgements – First and foremost, special thanks to Zsuzsanna Molnár for her enor- mous eff ort to sift through the relevant literature, as well as for her dedicated work in data entry and editing. Our active and retired colleagues in the Department of Palaeontology and Geology (István Főzy, Ottilia Szives, Attila Vörös, János Szabó) provided useful help in the collection under their care. Additional useful information on specimens and references were provided by Péter Ozsvárt, Attila Ősi, Piroska Pazonyi, Attila Virág, Matúš Hyžný, and Günther Schweigert. Access to the Mesozoic vertebrate collection was facilitated by Réka Kalmár. Eszter Hankó helped with data entry to the inventory.

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1. Triassic types 1.1. Triassic Radiolaria

Annulohaeckeliella curvispina Ozsvárt, Dumit- rica & Hungerbühler in Ozsvárt et al., 2017 holotype, PAL 2016.22.1.

Carnian, Killik Formation Tavusçayırı Tepe, Sorgun, Turkey Ozs17a, p. 154, pl. 3, fi g. 9

Archaeosemantis multispinosa Ozsvárt, Dumit- rica & Moix, 2017

holotype, PAL 2017.9.2.

Carnian, Killik Formation Tavusçayırı Tepe, Sorgun, Turkey Ozs17b, p. 65, fi g. 7.11

No inventory number given in original publication

paratype, PAL 2017.9.1.

Baumgartneria szarvaskoensis Ozsvárt &

Kovács, 2012

Ladinian, Extra-Bükkian olistoliths Malom-hegy, Szarvaskő

Ozs12, p. 281, pl. 2, fi gs 19–20

Kovács, 2012

Ozs12, p. 281, pl. 2, fi g. 21

Kovács, 2012

Ozs12, p. 281, pl. 2, fi g. 22

Capnuchosphaera barnabasi Kozur, Moix &

Ozsvárt, 2009 holotype, M 2009.1.1.

Carnian, Killik Formation Tavusçayırı Tepe, Sorgun, Turkey Koz09, p. 32, pl. 2, fi g. 1

Capnuchosphaera borbalae Kozur, Moix &

Capnuchosphaera bragini Kozur, Moix &

Capnuchosphaera ciliciensis Kozur, Moix &

Catalogue of invertebrate type specimens

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Capnuchosphaera crassa yforma Kozur, Moix &

Capnuchosphaera cylindrica cylindrica Kozur, Moix & Ozsvárt, 2009

holotype, M 2009.10.1.

Capnuchosphaera cylindrica retusaspinosa Kozur, Moix & Ozsvárt, 2009

Capnuchosphaera goestlingensis Kozur, Mostler

& Ozsvárt in Kozur et al., 2009 paratype, M 2009.12.1.

Carnian, Göstling Formation Göstling, Austria

Koz09, p. 34, pl. 2, fi g. 11

Capnuchosphaera gracilispinosa gracilispinosa Kozur, Moix & Ozsvárt, 2009

Capnuchosphaera gracilispinosa turkensis Kozur, Moix & Ozsvárt, 2009

Capnuchosphaera mersinensis Kozur, Moix &

Ozsvárt, 2009

Capnuchosphaera mostleri Kozur, Moix &

Ozsvárt, 2009

Capnuchosphaera multispinosa Kozur, Moix &

Ozsvárt, 2009

Capnuchosphaera oesii Kozur, Moix & Ozsvárt, 2009

Capnuchosphaera ottomanensis ottomanensis Kozur, Moix & Ozsvárt, 2009

Capnuchosphaera ottomanensis carterae Kozur, Moix & Ozsvárt, 2009

Capnuchosphaera tortuospinosa Kozur, Moix &

Ozsvárt, 2009

Capnuchosphaera tumorspinosa Kozur, Moix &

Ozsvárt, 2009

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Capnuchosphaera tuvalica Kozur, Moix &

Ozsvárt, 2009

Caponabolella brevispinosa Ozsvárt, Dumit rica

& Hungerbühler in Ozsvárt et al., 2017 holotype, PAL 2016.27.1.

Caponabolella brevispinosa Ozsvárt, Dumit rica

& Hungerbühler in Ozsvárt et al., 2017 paratype, PAL 2016.27.2.

Caponabolella longispinosa Ozsvárt, Dumit rica

Carnian, Killik Formation Tavusçayırı Tepe, Sorgun, Turkey Ozs17a, p. 156, pl. 5, fi gs 1–2

Caponabolella longispinosa Ozsvárt, Dumit ri ca

Circopoulpus cornubovis Ozsvárt, Dumit ri ca &

Hungerbühler in Ozsvárt et al., 2017 holotype, PAL 2016.21.1.

Circopoulpus cornubovis Ozsvárt, Dumitrica &

Hungerbühler in Ozsvárt et al., 2017 paratype, PAL 2016.21.3.

Circopoulpus cornubovis Ozsvárt, Dumitrica &

Circopoulpus cornubovis Ozsvárt, Dumit ri ca &

Circopoulpus dulaii Ozsvárt, Dumitrica &

Goricanella hexaspinosa Kozur, Mostler &

Ozsvárt in Kozur et al., 2009 holotype, M 2009.47.1.

Koz09, p. 43, pl. 4, fi g. 14

Ozsvárt in Kozur et al., 2009 paratype, M 2009.47.2.

Koz09, p. 43, pl. 4, fi g. 15

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Ozsvárt in Kozur et al., 2009 paratype, M 2009.47.3.

Koz09, p. 43, pl. 4, fi g. 16

Hindeosphaera burrii Ozsvárt, Moix & Kozur, 2015

Carnian, Killik Formation Tavusçayırı Tepe, Sorgun, Turkey Ozs15, p. 346, fi g. 5.23

Hindeosphaera djani Ozsvárt, Moix & Kozur, 2015

Hindeosphaera naomiae Ozsvárt, Moix &

Kozur, 2015

Minicrampus longispinosus Ozsvárt, Dumit rica

& Moix, 2017

Minicrampus longispinosus Ozsvárt, Dumit rica

& Moix, 2017

Monocoronella spinifera Ozsvárt, Dumitrica &

Moix, 2017

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Moix, 2017

Paramonocapnuchosphaera fusiformis Ozsvárt, Dumitrica & Moix, 2017

Paraweverella tenuispinosa Kozur, Moix &

Ozsvárt, 2009

Poulpus compactus Ozsvárt, Dumitrica & Hun- gerbühler in Ozsvárt et al., 2017

Poulpus elegans Ozsvárt, Dumitrica & Hunger- bühler in Ozsvárt et al., 2017

Poulpus kozuri Ozsvárt, Dumitrica & Hunger- bühler in Ozsvárt et al., 2017

Carnian, Killik Formation Tavusçayırı Tepe, Sorgun, Turkey Ozs17a, p. 145, pl. 1, fi gs 4–5

Pseudostylosphaera dumitricai Ozsvárt, Moix &

Kozur, 2015

Silicarmiger longospinosus Ozsvárt, Dumit rica

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Silicarmiger longospinosus Ozsvárt, Dumit rica

Spinocapnuchosphaera hantkeni Kozur, Moix &

Ozsvárt, 2009

Spinocapnuchosphaera odoghertyi Kozur, Moix

& Ozsvárt, 2009 holotype, M 2009.32.1.

Spinocapnuchosphaera szentei Kozur, Moix &

Ozsvárt, 2009

Spinocapnuchosphaera szivesae Kozur, Moix &

Ozsvárt, 2009

Spinocapnuchosphaera tekini tekini Kozur, Moix

& Ozsvárt, 2009 paratype, M 2009.28.2.

Spinocapnuchosphaera tekini hugluensis Kozur, Moix & Ozsvárt, 2009

Spinocapnuchosphaera tekini marginospinosa Kozur, Moix & Ozsvárt, 2009

Spinocapnuchosphaera tricuspidata Kozur, Moix

Spinocapnuchosphaera venusta Kozur, Moix &

Ozsvárt, 2009

Spinostylosphaera andrasi Ozsvárt, Moix &

Kozur, 2015

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Kozur, 2015

Carnian, Killik Formation Tavusçayırı Tepe, Sorgun, Turkey Ozs15, p. 349, fi gs 6.12.1–2

Inventory number given erroneously in fi gure caption as PAL 2014.127.3.6.

Spinostylosphaera masseti Ozsvárt, Dumit rica &

Moix, 2017

Spinostylosphaera masseti Ozsvárt, Dumit rica &

Moix, 2017

Spinostylosphaera michelae Ozsvárt, Dumit rica

& Moix, 2017

Spinostylosphaera sengoeri Ozsvárt, Dumit rica

& Moix, 2017

Spinostylosphaera vachardi Ozsvárt, Moix &

Kozur, 2015

Carnian, Killik Formation Tavusçayırı Tepe, Sorgun, Turkey Ozs15, p. 349, fi gs 6.13.1–2

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Kozur, 2015

Inventory number given erroneously in fi gure caption as PAL 2014.131.2.

Tamonella aspinosa Ozsvárt, Dumitrica &

Moix, 2017

Tamonella aspinosa Ozsvárt, Dumitrica &

Moix, 2017

Tekinium bragini Ozsvárt, Dumitrica &

Hungerbühler in Ozsvárt et al., 2017 holotype, PAL 2016.24.1.

Tekinium bragini Ozsvárt, Dumitrica &

Tetracapnuchosphaera? voeroesi Kozur, Moix &

Ozsvárt, 2009

Triassobullasphaera miriae Ozsvárt, Moix &

Kozur, 2015

Triassobullasphaera miriae Ozsvárt, Moix &

Kozur, 2015

Tubospongopallium inaequispinosum Kozur, Moix & Ozsvárt, 2009

Weverella gracilispinosa Kozur, Moix &

Ozsvárt, 2009

Weverella gracilispinosa Kozur, Moix &

Ozsvárt, 2009

paratype, M 2009.40.2.

Weverella longispinosa longispinosa Kozur, Moix

Weverella longispinosa curvata Kozur, Moix &

Ozsvárt, 2009

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Weverella longispinosa subrectangularis Kozur, Moix & Ozsvárt, 2009

Inventory number given erroneously in fi gure caption as M 2009.43.11.

1.2. Triassic Gastropoda Hungariella coronata Szabó, 2011 holotype, M 2010.427.1.

Norian–Rhaetian, Dachstein Limestone Formation

Remete-hegy, Budapest Sza11, p. 43, fi gs 2.5–8

Hungariella coronata Szabó, 2011 paratype, M 2010.425.1.

Remete-hegy, Budapest Sza11, p. 43, fi g. 2.12

Hungariella coronata Szabó, 2011 paratype, M 2010.428.1.

Remete-hegy, Budapest Sza11, p. 43, fi gs 2.9–11

Trochus (Mesotrochus) triadicus Kutassy, 1927 neotype, M 2010.422.1.

Remete-hegy, Budapest Kut27, p. 152 (pl. 5, fi gs 7a–c) Callotrochus triadicus, Sza11

Neotype designated and fi gured in Szabó 2011: p. 41, fi gs 1.1–3, 1.7–9

Trochus (Mesotrochus) triadicus elegantula Kutassy, 1933

holotype, M 2010.423.1.

Remete-hegy, Budapest Kut33, p. 236, pl. 2, fi g. 26 Callotrochus triadicus, Sza11

Type fi gured in Szabó 2011: fi gs 1.4–6

1.3. Triassic Cephalopoda

Epikellnerites pseudocholnokyi Vörös, 2018 holotype, PAL 2017.20.1.

Anisian, Vászoly Formation bed 9, section I, Mencshely VöA18, p. 71, pl. 9, fi gs 4a–b

Epikellnerites pseudocholnokyi Vörös, 2018 paratype, PAL 2017.21.1.

Anisian, Vászoly Formation Mencshely

VöA18, p. 71, pl. 9, fi gs 5a–b

VöA18, p. 71, pl. 9, fi gs 3a–b

VöA18, p. 71, text-fi gs 31–32 Epikellnerites spinatus Vörös, 2018 holotype, PAL 2017.23.1.

Anisian, Vászoly Formation bed 9, section I, Mencshely

VöA18, p. 72, text-fi g. 33, pl. 9, fi gs 6a–b Epikellnerites spinatus Vörös, 2018 paratype, PAL 2017.24.1.

VöA18, p. 72, pl. 9, fi g. 7 Epikellnerites tamasi Vörös, 2018 holotype, PAL 2017.10.1.

Anisian, Vászoly Formation?

vineyards, Szentbékkálla

VöA18, p. 67, text-fi gs 26–27, pl. 7, fi gs 3a–b Epikellnerites vaszolyensis Vörös, 2018 holotype, PAL 2017.12.1.

Anisian, Vászoly Formation trench P-11/c, Vászoly VöA18, p. 69, pl. 7, fi gs 6a–b

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Epikellnerites vaszolyensis Vörös, 2018 paratype, M.87.6

Anisian, Vászoly Formation?

Szentbékkálla

VöA18, p. 69, text-fi g. 30, pl. 8, fi gs 5a–b Epikellnerites vaszolyensis Vörös, 2018 paratype, M.98.14

Anisian, Vászoly Formation Vászoly

VöA18, p. 69, pl. 8, fi gs 2a–b

Epikellnerites vaszolyensis Vörös, 2018 paratype, M.98.74

VöA18, p. 69, text-fi g. 28c, pl. 8, fi gs 4a–b Epikellnerites vaszolyensis Vörös, 2018 paratype, PAL 2017.13.1.

Anisian, Vászoly Formation trench P-11/c, Vászoly

VöA18, p. 69, text-fi g. 28b, pl. 8, fi gs 1a–b Epikellnerites vaszolyensis Vörös, 2018 paratype, PAL 2017.14.1.

Anisian, Vászoly Formation trench P-11/c, Vászoly

VöA18, p. 69, text-fi g. 29, pl. 8, fi gs 3a–b Epikellnerites vaszolyensis Vörös, 2018 paratype, PAL 2017.15.1.

Epikellnerites vaszolyensis Vörös, 2018 paratype, PAL 2017.16.1.

Anisian, Vászoly Formation shaft P-XVIII, Vászoly VöA18, p. 69, pl. 8, fi gs 7a–b

VöA18, p. 69, pl. 8, fi gs 8a–b

Anisian, Vászoly Formation shaft P-XVIII, Vászoly VöA18, p. 69, pl. 9, fi gs 1a–b

VöA18, p. 69, text-fi g. 28a Hungarites sinuosus Vörös, 2018 holotype, PAL 2017.38.1.

VöA18, p. 120, text-fi gs 70a, 71, pl. 34, fi gs 5a–b Hungarites sinuosus Vörös, 2018

paratype, M.89.100 Anisian, Vászoly Formation bed 16/A, trench P-11/a, Vászoly

VöA18, p. 120, text-fi g. 70b, pl. 34, fi gs 4a–b Hungarites sinuosus Vörös, 2018

VöA18, p. 120, pl. 35, fi gs 1a–b Hungarites sinuosus Vörös, 2018 paratype, PAL 2017.40.1.

Anisian, Vászoly Formation Section I, Mencshely

VöA18, p. 120, pl. 35, fi gs 2a–b Hungarites sinuosus Vörös, 2018 paratype, PAL 2017.41.1.

VöA18, p. 120, pl. 35, fi gs 3a–b

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Hungarites szentei Vörös, 2018 holotype, PAL 2017.42.1.

Anisian, Vászoly Formation trench P-14, Vászoly

VöA18, p. 121, text-fi g. 72, pl. 36, fi gs 1a–b Hungarites szentei Vörös, 2018

VöA18, p. 121, pl. 36, fi gs 2a–b Hungarites szentei Vörös, 2018 paratype, PAL 2017.44.1.

Anisian, Vászoly Formation bed 6, Szentkirályszabadja VöA18, p. 121, pl. 36, fi gs 3a–b Hungarites szentei Vörös, 2018 paratype, PAL 2017.45.1.

Anisian, Vászoly Formation bed 5, Szentkirályszabadja VöA18, p. 121, pl. 36, fi g. 4 Hyparpadites szaboi Vörös, 2018 holotype, PAL 2017.25.1.

Anisian, Vászoly Formation shaft P-XVII, Vászoly

VöA18, p. 85, text-fi g. 43, pl. 11, fi gs 4a–b Hyparpadites szaboi Vörös, 2018

Anisian, Vászoly Formation shaft P-XVII, Vászoly VöA18, p. 85, pl. 11, fi gs 6a–b Hyparpadites szaboi Vörös, 2018 paratype, PAL 2017.27.1.

Anisian, Vászoly Formation bed 14, section I, Mencshely VöA18, p. 85, pl. 11, fi g. 3

Nodihungarites vinczei Vörös, 2018 holotype, M.89.81

Anisian, Vászoly Formation bed 16/A, trench P-11/a, Vászoly VöA18, p. 127, pl. 40, fi gs 1a–b

Nodihungarites vinczei Vörös, 2018 paratype, PAL 2017.46.1.

Anisian, Vászoly Formation trench P-11/a, Vászoly VöA18, p. 127, pl. 38, fi g. 7

Parahungarites solyensis Vörös, 2018 holotype, M.98.53

Anisian, Vászoly Formation bed 9, Őr-hegy, Sóly

VöA18, p. 98, pl. 18, fi gs 8a–b Parahungarites solyensis Vörös, 2018 paratype, M.98.208A

VöA18, p. 98, pl. 18, fi gs 5a–b Parahungarites solyensis Vörös, 2018 paratype, M.98.208B

VöA18, p. 98, text-fi g. 57, pl. 18, fi gs 9a–b Parahungarites solyensis Vörös, 2018 paratype, M.98.212A

VöA18, p. 98, pl. 18, fi gs 7a–b Parahungarites solyensis Vörös, 2018 paratype, PAL 2017.32.1.

VöA18, p. 98, pl. 18, fi gs 6a–b

Parakellnerites stuerzenbaumi Vörös, 2018 holotype, PAL 2017.30.1.

Anisian, Vászoly Formation trench P-14, Vászoly VöA18, p. 90, pl. 14, fi g. 1

Parakellnerites stuerzenbaumi Vörös, 2018 paratype, M.89.83

Anisian, Vászoly Formation bed 16/A, trench P-11/a, Vászoly VöA18, p. 90, pl. 14, fi gs 4a–b

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Parakellnerites stuerzenbaumi Vörös, 2018 paratype, M.89.86

Anisian, Vászoly Formation bed 16/A, trench P-11/a, Vászoly VöA18, p. 90, pl. 14, fi g. 3

Parakellnerites stuerzenbaumi Vörös, 2018 paratype, PAL 2017.28.1.

Anisian, Vászoly Formation bed 111/E, Felsőörs

VöA18, p. 90, pl. 13, fi gs 6a–b

Anisian, Vászoly Formation bed 111/F, Felsőörs VöA18, p. 90, pl. 13, fi g. 5

Anisian, Vászoly Formation bed 111/F, Felsőörs

VöA18, p. 90, pl. 14, fi gs 2a–b Stoppaniceras budaii Vörös, 2018 holotype, M.87.42

Anisian, Vászoly Formation bed 16/A, trench P-11/a, Vászoly

VöA18, p. 107, text-fi gs 60–61, pl. 27, fi gs 3a–b Stoppaniceras budaii Vörös, 2018

Anisian, Vászoly Formation bed 111, Felsőörs

VöA18, p. 107, pl. 27, fi gs 2a–b Stoppaniceras hermanni Vörös, 2018 holotype, M.98.16

Anisian, Vászoly Formation bed 16/A, trench P-11/a, Vászoly VöA18, p. 106, pl. 26, fi gs 5a–b Stoppaniceras hermanni Vörös, 2018 paratype, PAL 2017.36.1.

VöA18, p. 106, pl. 27, fi gs 1a–b

Stoppaniceras rieberi Vörös, 2018 holotype, M.98.26

VöA18, p. 103, pl. 25, fi gs 3a–b Stoppaniceras rieberi Vörös, 2018 paratype, M.98.27

VöA18, p. 103, pl. 25, fi g. 7 Stoppaniceras rieberi Vörös, 2018 paratype, PAL 2017.33.1.

Anisian, Vászoly Formation bed 111/I, Felsőörs

VöA18, p. 103, pl. 25, fi gs 4a–b Stoppaniceras rieberi Vörös, 2018 paratype, PAL 2017.34.1.

VöA18, p. 103, pl. 25, fi g. 6 Stoppaniceras rieberi Vörös, 2018 paratype, PAL 2017.35.1.

VöA18, p. 103, pl. 25, fi g. 5 2. Jurassic types

2.1. Jurassic Bivalvia

Pholadomya schafarziki Papp, 1907 holotype, M.63.3198

Callovian–Oxfordian, “limestone”

Gunib, Daghestan, Russia Pap07, p. 155, pl. 3, fi gs 1, 1a–b

Pholadomya (Bucardiomya) lirata, Gal08 Type fi gured in Galácz & Szente 2008: pl.

4, fi gs 1–4

Pleuromya merzbacheri Papp, 1907 holotype, M.63.3616

Callovian

Psebay, Kuban, Russia

Pap07, p. 158, pl. 4, fi gs 2, 2a–c Pleuromya alduini, Gal08

Type fi gured in Galácz & Szente 2008: pl.

4, fi gs 8–9

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2.2. Jurassic Gastropoda Adeorbisina procera Szabó, 1981

paratypes, PAL 2018.34.1., PAL 2018.35.1., PAL 2018.36.1–4.

Bajocian, fi ssure-fi lling of Csókakő Limestone Formation?

Som-hegy, Bakonybél Sza81, p. 63

Mariottia gibbosa, Con87 Anoptychia hastata Szabó, 1983 paratype, PAL 2018.37.1.

Bakonyia planapex Szabó, 1981

paratypes, PAL 2018.38.1–2., PAL 2018.39.1–

2., PAL 2018.40.1–2.

Bathrotomaria mandokii Szabó, 1980 paratype, PAL 2018.41.1.

Dimorphotectus unicarinatus Szabó, 1981 paratypes, PAL 2018.42.1–3., PAL 2018.43.3–4.

Eucycloidea galaczi Szabó, 1983

paratypes, PAL 2018.43.5., PAL 2018.45.1–3., PAL 2018.46.1–2., PAL 2018.47.1., PAL 2018.48.1.

Eucycloidea galaczi Szabó, 1983 paratype, PAL 2018.44.1.

Som-hegy, Bakonybél Sza83, p. 42, pl. 3, fi g. 8 Laevitomaria danii Szabó, 2009 holotype, M 2008.542.1.

Pliensbachian, Hierlatz Limestone Formation Fenyveskút, Lókút

Sza09, p. 49, fi gs 43a–c

Leptomaria somhegyensis Szabó, 1980 paratype, PAL 2018.49.1.

Som-hegy, Bakonybél Sza80, p. 59, pl. 3, fi g. 2

Trochotomaria somhegyensis, Con87 Leptomaria somhegyensis Szabó, 1980 paratypes, PAL 2018.50.1–4.

Trochotomaria somhegyensis, Con87 Neritopsis (Neritopsis) spinigera Szabó, 1982 paratypes, PAL 2018.51.1., PAL 2018.52.1., PAL 2018.54.1–6., PAL 2018.58.1.

Neritopsis (Neritopsis) spinigera Szabó, 1982 paratypes, PAL 2018.53.1–17.

Som-hegy, Bakonybél Sza82, p. 19, pl. 1, fi g. 12

Neritopsis (Neritopsis) spinigera Szabó, 1982 paratype, PAL 2018.55.1.

Som-hegy, Bakonybél Sza82, p. 19, pl. 1, fi gs 15–16

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Som-hegy, Bakonybél Sza82, p. 19, pl. 1, fi gs 17–18 Ochetochilus piceus Szabó, 1983 paratypes, PAL 2018.59.1–2.

Som-hegy, Bakonybél Sza83, p. 43, pl. 3, fi gs 1–3 Telleria (Telleria) picea, Con87 Pietteia trispinigera Szabó, 1983

paratypes, PAL 2018.60.1–2., PAL 2018.61.1–2.

Pietteia (Trietteia) trispinigera, Con87 Procerithium? (Cosmocerithium?) angulocos- tatum Szabó, 1983

paratypes, PAL 2018.62.1–2.

Diatrypesis angulocostatum, Con87 Proconulus epuliformis Szabó, 1981 paratype, PAL 2018.63.1.

Type fi gured in Conti & Szabó 1987: fi g. 1e

Proconulus epuliformis Szabó, 1981

paratypes, PAL 2018.64.1–13., PAL 2018.65.1., PAL 2018.66.1–11., PAL 2018.67.1–7., PAL 2018.68.1–2., PAL 2018.69.1–4.

Proconulus rimosus Szabó, 1981

paratypes, PAL 2018.70.1–2., PAL 2018.71.1–

4., PAL 2018.72.1., PAL 2018.73.1–13., PAL 2018.74.1., PAL 2018.75.1., PAL 2018.76.1–11., PAL 2018.79.1–3.

Proconulus baldensis, Con87 Proconulus rimosus Szabó, 1981 paratypes, PAL 2018.77.1–6.

Proconulus baldensis, Con87

One of the paratypes fi gured in Conti &

Szabó 1987: fi g. 2

Proconulus rimosus Szabó, 1981 paratype, PAL 2018.78.1.

Proconulus baldensis, Con87

Type fi gured in Conti & Szabó 1987: fi g. 2 Pyrgotrochus? problematicus Szabó, 1980 paratype, PAL 2018.80.1.

Laevitomaria problematica, Con87 Type fi gured under inventory number INV.2012.15.1 in Gatto et al. 2015: fi g. 2c_1–3

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Pyrgotrochus? problematicus Szabó, 1980 paratype, PAL 2018.80.2.

Laevitomaria problematica, Con87 Type fi gured under inventory number INV.2012.15.2 in Gatto et al. 2015: fi g. 2d_1–3 Pyrgotrochus? problematicus Szabó, 1980 paratype, PAL 2018.80.3.

Laevitomaria problematica, Con87 Type fi gured under inventory number INV.2012.15.3 in Gatto et al. 2015: fi g. 2b Pyrgotrochus? problematicus Szabó, 1980 paratypes, PAL 2018.80.4–7., PAL 2018.81.1–

6., PAL 2018.82.1., PAL 2018.83.1.

Laevitomaria problematica, Con87 Trochopsidea kondai Szabó, 1981

paratypes, PAL 2018.84.1., PAL 2018.85.1–4., PAL 2018.86.1–11., PAL 2018.87.1–5.

Ventricaria? vesicula Szabó, 1983 paratypes, PAL 2018.88.1–2.

Som-hegy, Bakonybél Sza83, p. 30, pl. 1, fi g. 10 Ventricaria? vesicula Szabó, 1983 paratypes, PAL 2018.89.1–2.

Zircia zircensis Szabó, 1981

paratypes, PAL 2018.90.1–2., PAL 2018.91.1., PAL 2018.92.1., PAL 2018.93.1.

2.3. Jurassic Cephalopoda

Perisphinctes daghestanicus Papp, 1907 holotype, M.63.3304A

Bajocian, “lower dark shale”

Gunib, Daghestan, Russia Pap07, p. 166, pl. 7, fi gs 3, 3a Prorsisphinctes loczyi, Gal08

3, fi g. 7, specimen found in collection in 2008 Perisphinctes lóczyi Papp, 1907

lectotype, M.63.3301A Bajocian, “lower dark shale”

Gunib, Daghestan, Russia Pap07, p. 165, pl. 7, fi g. 1 Prorsisphinctes loczyi, Gal08

Lectotype designated and fi gured in Galácz

& Szente 2008: p. 114, pl. 3, fi gs 1–2 Perisphinctes lóczyi Papp, 1907 paralectotype, M.63.3301B Bajocian, “lower dark shale”

Gunib, Daghestan, Russia Pap07, p. 165, pl. 7, fi gs 2, 2a–b Prorsisphinctes loczyi, Gal08

Paralectotype No. 1 fi gured in Galácz &

Szente 2008: pl. 3, fi gs 5–6 Perisphinctes lóczyi Papp, 1907 paralectotype, M.63.3208 Bajocian, “lower dark shale”

Gunib, Daghestan, Russia Pap07, p. 165

Prorsisphinctes loczyi, Gal08

Counterpart of paralectotype No. 1, specimen found in collection in 2008

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Perisphinctes lóczyi Papp, 1907 paralectotype, M.63.3304A Bajocian, “lower dark shale”

Paralectotype No. 2 fi gured in Galácz &

Szente 2008: pl. 3, fi g. 7, specimen found in collection in 2008

Perisphinctes lóczyi Papp, 1907 paralectotype, M.63.3304B Bajocian, “lower dark shale”

Paralectotype No. 3, specimen found in collection in 2008

Simoceras agostyani Főzy & Scherzinger, 2013 holotype, PAL 2013.26.1.

Tithonian, Szentivánhegy Limestone Formation

bed 12, section I, Szomód Főz13, p. 254, pl. 27, fi gs 1a–b

Original species name emended here as agosty- anense

Simoceras szentei Főzy & Scherzinger, 2011 holotype, PAL 2011.4.1.

Tithonian, Szentivánhegy Limestone Formation

bed 5, Paprét-árok, Süttő Főz11, p. 124, fi gs 5.1, 6.2

Stephanoceras liechtensteinii Papp, 1907 holotype, M.63.3610

Bajocian, “dark ferruginous limestone boul- der”

Fiagdon Creek, west of Vladikavkaz, Russia Pap07, p. 163, pl. 6, fi g. 4

2, fi gs 5–6

Virgatosimoceras dunaii Scherzinger, Főzy &

Parent, 2010 holotype, M.92.749

Tithonian, Pálihálás Limestone Formation bed 47, Lókúti-domb, Lókút

ScA10, p. 201, fi gs 4.1a–b

2.4. Jurassic Brachiopoda Apringia barnabasi Vörös, 2009 holotype, 2007.121.1.

Pliensbachian, Hierlatz Limestone Formation bed 26, Kericser section, Lókút

VöA09, p. 45, pl. 1, fi gs 6a–c Apringia barnabasi Vörös, 2009 paratype, 2007.122.1.

VöA09, p. 45, pl. 1, fi gs 7a–c Cirpa alkayae Vörös, 2014 holotype, PAL 2014.137.1.

Pliensbachian, Bayırköy Formation Beytepe (Lodumlu), Ankara area, Turkey VöA14b, p. 19, fi gs 17–20

Cirpa alkayae Vörös, 2014 paratype, PAL 2014.138.1.

Pliensbachian, Bayırköy Formation Beytepe (Lodumlu), Ankara area, Turkey VöA14b, p. 19, fi g. 23

Fenyveskutella pseudouhligi Vörös, 2009 holotype, 2007.229.1.

Pliensbachian, Tűzkövesárok Limestone Formation

basal layer, Fg-I, Gombáspuszta, Szentgál VöA09, p. 90, pl. 10, fi gs 9a–c

Fenyveskutella pseudouhligi Vörös, 2009 paratype, 2007.228.1.

Pliensbachian, Hierlatz Limestone Formation Kericser section, Lókút

VöA09, p. 90, pl. 10, fi gs 8a–c Fenyveskutella theresiae Vörös, 2009 holotype, 2007.231.1.

Pliensbachian, Hierlatz Limestone Formation grab sample É, Fenyveskút, Lókút

VöA09, p. 91, pl. 10, fi gs 11a–c Fenyveskutella vighi Vörös, 2009 holotype, 2007.221.1.

Pliensbachian, Hierlatz Limestone Formation grab sample p1, Fenyveskút, Lókút

VöA09, p. 88, pl. 10, fi gs 1a–c

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Fenyveskutella vighi Vörös, 2009 paratype, 2007.222.1.

Pliensbachian, Hierlatz Limestone Formation grab sample É, Fenyveskút, Lókút

VöA09, p. 88, pl. 10, fi gs 2a–c Karadagella szentei Vörös, 2013 holotype, PAL 2013.1.1.

Oxfordian, Pálihálás Limestone Formation?

Velka Skala, Kétágú-hegy, Kesztölc VöA13, p. 384, pl. 1, fi gs 6a–c Karadagella szentei Vörös, 2013 paratype, PAL 2013.2.1.

Oxfordian, Pálihálás Limestone Formation?

Velka Skala, Kétágú-hegy, Kesztölc VöA13, p. 384, text-fi g. 4

Paronarhynchia estherae Vörös, 2009 holotype, 2007.237.1.

VöA09, p. 101, pl. 11, fi gs 1a–c Paronarhynchia estherae Vörös, 2009 paratype, 2007.239.1.

VöA09, p. 101, pl. 11, fi gs 3a–c

Prionorhynchia? catharinae Vörös, 2009 holotype, 2007.192.1.

Pliensbachian, Hierlatz Limestone Formation grab sample D, Fenyveskút, Lókút

VöA09, p. 70, pl. 8, fi gs 6a–c Rhapidothyris lokutica Vörös, 2009 holotype, M 2008.391.1.

VöA09, p. 145, pl. 16, fi gs 4a–c Rhapidothyris lokutica Vörös, 2009 paratype, M 2008.390.1.

Pliensbachian, Isztimér Limestone Formation bed 3, Büdöskút, Gyulafi rátót, Veszprém VöA09, p. 145, pl. 16, fi gs 3a–c

3. Cretaceous types

3.1. Cretaceous Cephalopoda Ancyloceras capellini Matheron, 1880 holotype, M.62.3

Barremian Clumanc, France

Mat80, pl. C-24, fi gs 1a–b Heteroceras? capellini, Kle07

Specimen previously in permanent exhibition of the museum, returned to collection in 2013

?Chigaroceras szomodi Szives & Főzy, 2013 holotype, PAL 2013.29.1.

Berriasian, Szentivánhegy Limestone Formation

bed 10, section I, Szomód Szi13, p. 319, pl. 6, fi gs 1a–b

Original species name emended here as szo- modense

3.2. Cretaceous Brachiopoda Sphenope bifi da Vörös, 2014 holotype, PAL 2013.27.1.

bed 41, Szilas-árok, Borzavár VöA14a, p. 9, fi gs 26–28 Sphenope bifi da Vörös, 2014 paratype, PAL 2013.28.1.

bed 41, Szilas-árok, Borzavár VöA14a, p. 9, fi g. 36

3.3. Cretaceous Crinoidea

Apsidocrinus doreckae Konieczyński, Pisera &

Főzy, 2016

Barremian, Bersek Marl Formation bed 252, Bersek-hegy, Lábatlan KPF16, p. 162, fi gs 4l–m

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Apsidocrinus doreckae Konieczyński, Pisera &

Főzy, 2016

Barremian, Bersek Marl Formation bed 252, Bersek-hegy, Lábatlan KPF16, p. 162, fi g. 4n

Phyllocrinus bersekensis Konieczyński, Pisera &

Főzy, 2016

Barremian, Bersek Marl Formation bed 257, Bersek-hegy, Lábatlan KPF16, p. 157, fi gs 3k–l

Phyllocrinus bersekensis Konieczyński, Pisera &

Főzy, 2016

Barremian, Bersek Marl Formation bed 258, Bersek-hegy, Lábatlan KPF16, p. 157, fi g. 3i

3.4. Cretaceous Echinoidea

Absurdaster hungaricus Kroh, Lukeneder &

Gallemí, 2014

Hauterivian, Bersek Marl Formation bed 221, Bersek-hegy, Lábatlan

Kro14, p. 244, fi gs 5.3, 6.3a–b, 7.1a–c, 8.1a–b Absurdaster hungaricus Kroh, Lukeneder &

Gallemí, 2014

Hauterivian, Bersek Marl Formation bed 226, Bersek-hegy, Lábatlan Kro14, p. 244, fi gs 7.2, 8.2

Absurdaster hungaricus Kroh, Lukeneder &

Gallemí, 2014

Hauterivian, Bersek Marl Formation bed 228, Bersek-hegy, Lábatlan

Kro14, p. 244, fi gs 5.4, 6.4a–b, 7.3a–c, 8.3 Absurdaster hungaricus Kroh, Lukeneder &

Gallemí, 2014

Hauterivian, Bersek Marl Formation bed 233, Bersek-hegy, Lábatlan Kro14, p. 244, fi gs 7.4a–b, 9a–c

4. Eocene types

4.1. Eocene Foraminiferida

Globigerina applanata Hantken, 1883 lectotype, M.72.553

Priabonian, “marl”

l’Escarène (Scarena), France

Han83a (Han83b), p. 132 (11), pl. 2, fi gs 7a–c Turborotalia applanata, Szt73

Lectotype designated and fi gured in Sztrákos 1973: p. 224, pl. 1, fi gs 3a–c. Type found in collection in 2009

Globigerina applanata Hantken, 1883 paralectotype, M.72.554

l’Escarène (Scarena), France Han83a (Han83b), p. 132 (11) Turborotalia applanata, Szt73

Type fi gured in Sztrákos 1973: pl. 1, fi gs 4a–c. Type found in collection in 2009 Globigerina applanata Hantken, 1883 paralectotype, M.72.555

l’Escarène (Scarena), France Han83a (Han83b), p. 132 (11) Turborotalia applanata, Szt73

Type fi gured in Sztrákos 1973: pl. 1, fi gs 5a–c. Type found in collection in 2009 Globigerina applanata Hantken, 1883

paralectotypes, M.72.556, M.72.557, M.72.558 Priabonian, “marl”

l’Escarène (Scarena), France Han83a (Han83b), p. 132 (11) Turborotalia applanata, Szt73 Types found in collection in 2009 Globigerina globosa Hantken, 1883 neotype, M.72.548

l’Escarène (Scarena), France

Han83a (Han83b), p. 132 (11) (pl. 2, fi gs 3a–b) Globigerinatheca globosa, Szt73

Neotype designated and fi gured in Sztrákos 1973: p. 228, pl. 2, fi gs 1a–c. Type found in collection in 2009

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4.2. Eocene Polyplacophora

Lepidochitona viciani Dell’Angelo, Sosso, Kroh

& Dulai, 2015

Lutetian–Bartonian, Kincses Formation roadcut at the vineries, Gánt

Del15, p. 363 Head valve

& Dulai, 2015

Del15, p. 363 Intermediate valve

& Dulai, 2015

paratype, PAL 2013.36.1

Del15, p. 363, fi gs 5g–i, 6d Tail valve

4.3. Eocene Brachiopoda

Magellania hantkeni Meznerics, 1944 neotype, M 2009.568.1.

Lutetian, Szőc Limestone Formation Öreg-hegy quarry, Csojjányos-völgy, Kislőd Mez44, p. 46 (pl. 3, fi gs 13–16, pl. 5, fi gs 21–23)

Meznericsia hantkeni, Bit11

Neotype designated and fi gured in Bitner et al. 2011: p. 121, fi gs 6g–i

Rugia zagorseki Dulai, 2011 holotype, M 2010.281.1.

Priabonian, Perwang Formation 2377.5 m, borehole Perwang-1, Austria Dul11a, p. 302, fi gs 6e–f

Rugia zagorseki Dulai, 2011 paratype, M 2010.278.1.

Priabonian, Perwang Formation 3194 m, borehole Helmberg-1, Austria Dul11a, p. 302, fi g. 6a

Priabonian, Perwang Formation 3194 m, borehole Helmberg-1, Austria Dul11a, p. 302, fi gs 6b–c

Priabonian, Perwang Formation 2083.5 m, borehole Perwang-1, Austria Dul11a, p. 302, fi g. 6d

Terebratulina johansenae Dulai, 2011 holotype, M 2010.275.1.

Priabonian, Perwang Formation 2073 m, borehole Perwang-1, Austria Dul11a, p. 300, fi g. 5f

Terebratulina johansenae Dulai, 2011 paratype, M 2010.270.1.

Priabonian, Perwang Formation 2054.5 m, borehole Perwang-1, Austria Dul11a, p. 300, fi g. 5a

Priabonian, Perwang Formation 2054.5 m, borehole Perwang-1, Austria Dul11a, p. 300, fi g. 5b

Priabonian, Perwang Formation 2073 m, borehole Perwang-1, Austria Dul11a, p. 300, fi g. 5c

Priabonian, Perwang Formation 2073 m, borehole Perwang-1, Austria Dul11a, p. 300, fi g. 5d

Priabonian, Perwang Formation 2073 m, borehole Perwang-1, Austria Dul11a, p. 300, fi g. 5e

FRAGMENTA PALAEONTOLOGICA HUNGARICAVolume 35 Budapest, 2018 pp. 3–59

Volume 35 Budapest, 2018 pp. 3–59

First supplement to the catalogue of

invertebrate and vertebrate palaeontological type specimens of the Hungarian Natural History Museum: 2008–2018

TABLE OF CONTENTS

Introduction 5

Catalogue of invertebrate type specimens 10

1. Triassic types 10

1.1. Triassic Radiolaria 10

1.2. Triassic Gastropoda 18

1.3. Triassic Cephalopoda 18

2. Jurassic types 21

2.1. Jurassic Bivalvia 21

2.2. Jurassic Gastropoda 22

2.3. Jurassic Cephalopoda 24

2.4. Jurassic Brachiopoda 25

3. Cretaceous types 26

3.1. Cretaceous Cephalopoda 26

3.2. Cretaceous Brachiopoda 26

3.3. Cretaceous Crinoidea 26

3.4. Cretaceous Echinoidea 27

4. Eocene types 27

4.1. Eocene Foraminiferida 27

4.2. Eocene Polyplacophora 28

4.3. Eocene Brachiopoda 28

5. Oligocene types 29

5.1. Oligocene Gastropoda 29

5.2. Oligocene Decapoda 29

6. Miocene types 32

6.1. Miocene Bivalvia 32

6.2. Miocene Gastropoda 32

6.3. Miocene Brachiopoda 34

6.4. Miocene Decapoda 34

6.5. Miocene Echinoidea 34

Catalogue of vertebrate type specimens 35

7. Amphibia 35

8. Reptilia 36

9. Aves 46

10. Mammalia 46

Index of species and subspecies names 49

References 52

INTRODUCTION

Aft er moving of the palaeontological collections to a new location in 2004–

Th e catalogue provided a comprehensive listing of invertebrate and vertebrate type specimens published between 1841 and February 2008. Th e palaeobotani- cal type specimens are historically part of the collection of the Department of Botany at the HNHM (Hably & Szakály 1989; Hably et al. 2001).

Several type specimens were recently discovered in some historical collections or older materials, or were donated to the HNHM (e.g., Papp 1907; Kutassy 1933;

Somos & Kókay 1960; Steininger et al. 1973).

Following the structure of the original catalogue, this supplement also in-

cludes primary, name-bearing types (holotypes, syntypes, lectotypes, neotypes)

and secondary types (paratypes, paralectotypes). It is subdivided into separate

parts for invertebrate and vertebrate types. Th e basic information of taxa and

their type specimens is listed in slightly diff erent formats for invertebrates and

vertebrates. An individual catalogue entry typically corresponds to an invento-

ry lot that contains the type specimen(s) of a taxon, identifi ed by an inventory

number. If a taxon is represented by more than one type specimen, separate en-

tries are listed where diff erences exist in either the inventory number, locality,

or illustration of specimen. Types under more than one inventory number are

merged into a single entry if all data fi elds other than the inventory number are

identical.

Th e format for invertebrate entries is as follows:

Th e format for pre-Pliocene vertebrate entries is as follows:

Th e format for Pliocene and Quaternary vertebrate entries is as follows:

Th e invertebrate catalogue entries are arranged primarily by geologic age, secondarily by major taxonomic groups, and then the original proposed bi- or trinomens are listed alphabetically within each subdivision.

Secondary, taxonomic subdivisions use the following, well-known groups, arbi- trarily taken from higher taxa at the phylum, class, or order level: Radiolaria, Foraminiferida, Polyplacophora, Bivalvia, Gastropoda, Cephalopoda, Brachio- poda, Decapoda, Crinoidea, Echinoidea.

Th e catalogue of vertebrate types is arranged by higher taxa, containing sub-

divisions for Amphibia, Reptilia, Aves, and Mammalia.

völgy – valley. Geological terms for localities, such as outcrop (“feltárás”), quarry

(“kőfejtő”), brickyard (“téglagyár”), coal mine (“szénbánya”) are given in English.

Name of town refers to the municipality to which the locality belongs, in some cases this is not the nearest settlement.

As the majority of the types were collected at localities within the present- day borders of Hungary (385 inventory lots), no country name is listed for them.

Illustrations (text-fi gures and/or fi gures on plates) accompanying the origi- nal description are quoted at the appropriate entry, if the specimen is identifi ed as the fi gured one.

If we became aware of a published revision, either nomenclatural, or changing the species-level identifi cation or the generic assignment of the taxon concerned, we include the revised name and provide the reference code of the revising work.

Any additional information may appear as remarks in the last line of an item.

Remarks may include reference to a diff erent inventory number in the original pub- lication, notes on the type designation, or the date of rediscovery in the collection.

Although signifi cant care was taken to compile a comprehensive catalogue

that contains accurate information, errors and omissions are almost inevitable in

an endeavour of this scale. Th erefore users of this catalogue are encouraged to re-

port any suggested correction and/or addition to the curators of the Department

of Palaeontology and Geology.

*

Catalogue of invertebrate type specimens