sect.
Canescentia
ser. Eladia sect. Eladia
0.1
P. chroogomphum P. americanum P. lenticrescens P. soppii
91/*
P. tunisiense P. lusitanum P. virgatum
87/*
P. brevicompactum P. kongii
P. neocrassum P. fennelliae P. bialowiezense P. astrolabium
P. olsonii P. salamii
94/*
P. spathulatum P. buchwaldii P. tularense
85/-P. arizonense P. yarmokense P. canescens P. radiatolobatum P. murcianum P. jensenii
P. janczewskii P. dunedinense
93/*
P. nigricans P. griseoazureum P. corvianum
“P. ochotense” (=P. antarcticum)
“P. attenuatum” (=P. antarcticum) P. antarcticum
“P. piltunense” (=P. antarcticum)
84
-P. atrovenetum P. coralligerum
P. nucicola
79/*
91/*
P. novae-zeelandiae P. sacculum
P. senticosum A. glaucus
H. avellanea
-/0.95
-/0.95
-/0.98
-/*
C
Fig. 19. (Continued).
Tal.avellaneus), but differs in ascomatal initials, ascospore ornamen-tation and conidiophore branching (Kong 1999).Samsonet al.(2011c) indicated that Tal. brevicompactus represents a distinct species in Hamigera and our phylogenetic analysis (Supplementary Fig. S2) supports this observation. The new combinationH. brevicompacta is proposed here.
Paecilomyces lagunculariae(C. Ram) Houbraken, Frisvad & Samson, comb.nov.MycoBank MB832559.
Basionym: Byssochlamys nivea var. lagunculariae C. Ram, Nova Hedwigia 16: 311. 1968.
Synonym:Byssochlamys lagunculariae (C. Ram) Samsonet al., Per-soonia 22: 18. 2009.
Notes:Ram (1968)describedByssochlamys niveavar.lagunculariae, without the description of thePaecilomycesasexual morph. In a later study on the taxonomy ofByssochlamysand itsPaecilomycesasexual morphs,Samsonet al.(2009)elevated this variety to species level, but also did not formally describe thePaecilomycesmorph. The type spe-cies of Paecilomyces, Paec. variotii, and the type species of Byssochlamys,B. nivea, were shown to be congeneric through mo-lecular sequence analyses (Luangsa-Ardet al. 2004, Samson et al.
2009). Anticipating on the change to one scientific name for fungi (McNeillet al.2012),Rossmanet al.(2016)recommended the use of PaecilomycesoverByssochlamys, and we therefore formally introduce Paec.lagunculariaehere.
Penicillago kabunica(Baghd.) Houbraken, Frisvad & Samson,comb.
nov.MycoBank MB832560.
Basionym:Penicillium kabunicumBaghd., Novosti Sist. Nizsh. Rast. 5:
98. 1968.
Notes: See underPenicillago.Penicillium kabunicum(andP. moldavi-cum) were treated byRamírez (1982)in the“P.brasilianum-series”and Pitt (1980)treated this species as a synonym ofP.janthinellum. These observations indicate a relationship with Penicillium sect. Lanata-Divaricata; however, phylogenetic analysis does not support this placement.Penicillago kabunicaproduces cycloaspeptide A and D.
Penicillago mirabilis (Beliakova & Milko) Houbraken, Frisvad &
Samson,comb.nov.MycoBank MB832561.
Basionym:Penicillium mirabileBeliakova & Milko, Mikol. Fitopatol. 6:
145. 1972.
Notes: See also above, underPenicillago(under genera). The identity of this species is controversial and needs further study. Samson et al.
(2011c) andYilmaz et al. (2014) showed in their phylogenetic ana-lyses that P. mirabile belongs to Talaromyces sect. Trachyspermi.
However,Samson et al.(2011c)had doubts about these results and they therefore did not combine this species in Talaromyces. Further-more, they noted that the type strain is in poor condition. Two sequences obtained from IMI 167383 (ex-type ofP.mirabile) (KC992257,BenA;
KC962096, ITS) are present in GenBank and comparison of those sequences indicate that this strain is aPenicillium corylophilum. In the original description ofP.mirabile, it is mentioned that the species pro-duces echinulate conidia with conspicuous connectives. This feature does notfit withP.corylophilum and the sequences derived from IMI 167383 are incorrect.Pitt (1980)placedP.mirabileinPenicillium sub-gen.Biverticillium(nowadaysTalaromyces pro parte). However, he also reported production of smooth-walled conidia and the strain he exam-ined therefore also does not fit with the original description. Re-examination of CBS 624.72, the ex-type of P. mirabile, shows that this strainfits well with the description ofRamírez (1982). Similar to the observation ofRamírez (1982), deep brown to black coloured sclerotia were not observed in CBS 624.72. Phylogenetic analysis usingBenA, CaM, ITS andRPB2sequences shows that this strain is related toP. kabunicum,P.moldavicumandP.nodositatumin the genusPenicillago (Fig. 6).
Penicillago moldavica (Milko & Beliakova) Houbraken, Frisvad &
Samson,comb.nov.MycoBank MB832562.
Basionym: Penicillium moldavicum Milko & Beliakova, Novosti Sist.
Nizsh. Rast. 4: 255. 1967.
Notes: See above, under Penicillago, and in Guevara-Suarez et al.
(2020).Penicillago moldavicaproduces alternatriol.
Penicillago nodositata (Valla) Guevara-Suarez et al., Fungal Syst.
Evol. 5: 64. 2020. MycoBank MB822074.
Basionym:Penicillium nodositatumValla, Pl. & Soil 114: 146. 1989.
Notes: Based on sequence data,P.nodositatumwas tentatively placed in synonymy with P. bilaiae by Houbraken & Samson (2011). This conclusion was drawn based on incorrect accession numbers in the CBS culture collection (Visagieet al.2013).Visagieet al.(2013) sub-sequently re-examined CBS 333.90, the ex-type ofP.nodositatum, and reported that this strain is related toP.kabunicum, outsidePenicillium sensu stricto. Penicillago nodositatais unique in that it forms myco-nodules in roots ofAlnus trees (Valla et al. 1989). We detected an altersolanol, cycloaspeptide A and griseofulvin in this species.
Phialomyces arenicola (Chalab.) Houbraken, Frisvad & Samson, comb.nov.MycoBank MB832563.
Basionym: Penicillium arenicola Chalab., Not. Syst. Crypt. Inst. bot.
Acad. Sci. USSR: 162. 1950.
Synonym:Penicillium canadenseG. Sm., Trans. Brit. Mycol. Soc. 39:
113. 1956.
Notes: Merimbla humicoloides, P. arenicola and P. canadense are phylogenetically related toPhialomyces macrosporus, the type species ofPhialomyces(Fig. 1) (Misra & Talbot 1964). Based on a multigene phylogenetic analysis, M. humicoloides proved not to belong to the Hamigera-clade. To maintain a monophyletic genus, Peterson et al.
(2010)decided to combine this species in the genusPenicillium, as P. humicoloides. On the other hand, they also noted a close rela-tionship of this species with P. arenicola and P. canadensis, two species outsidePenicillium sensu stricto(Houbraken & Samson 2011).
The species belonging to thePhialomycesclade produce penicillium-like conidiophores and gold-brown conidia, a feature not observed in Penicillium sensu stricto. Based on phylogenetic data and morphology, we combineMerimbla(=Penicillium)humicoloides, andP.arenicolain Phialomyces. Pitt (1980) treated P. canadense as a synonym of P. arenicola. A phylogenetic analysis of six loci resolvedP.canadenseas a close relative of P. arenicola (Peterson et al. 2010) but since no phenotypic differences exist between these species we regard Peni-cillium canadense as a synonym of Phialomyces arenicola. Phialo-myces arenicola has been reported to produce canadensolides (McCorkindaleet al.1968), chlorogentisylalcohol (McCorkindaleet al.
1972) and asperphenamate (McCorkindaleet al.1978).
Phialomyces humicoloides (Bills & Heredia) Houbraken, Frisvad &
Samson,comb.nov.MycoBank MB832564.
Basionym: Merimbla humicoloides Bills & Heredia, Mycol. Res.105:
1276. 2001.
Synonym: Penicillium humicoloides (Bills & Heredia) S.W. Peterson et al., Mycologia 102: 858. 2010.
Note: SeePh.arenicola.
Pseudohamigera striata (Raper & Fennell) Houbraken, Frisvad &
Samson,comb.nov.MycoBank MB832565.
Basionym: Penicillium striatumRaper & Fennell, Mycologia 40: 521.
1948.
Synonyms:Talaromyces striatus (Raper & Fennell) C.R. Benj., Myco-logia 47: 682. 1955.
Hamigera striata(Raper & Fennell) Stolk & Samson, Persoonia 6: 347.
1971.
Byssochlamys striata(Raper & Fennell) Arx, Mycotaxon 26: 120. 1986.
Penicillium lineatumPitt, The Genus Penicillium: 485. 1980.
Paecilomyces lineatus(Pitt) Arx, Mycotaxon 26: 120. 1986.
Notes: The taxonomic position of Pseudohamigera striata was dis-cussed several times in history. The sexual morph ofPseudohamigera
striata was previously classified in Talaromyces, Hamigera and Byssochlamysand the asexual morph inPenicilliumandPaecilomyces (Raper & Fennell 1948, Stolk & Samson 1971, Pitt 1980, von Arx 1986), showing the difficulty in classifying this species. Phylogenetic data (Supplementary Fig. S1) show that this species is distinct from Byssochlamys,Hamigera,Paecilomyces,PenicilliumandTalaromyces.
Talaromyces resinae(Z.T. Qi & H.Z. Kong) Houbraken & X.C. Wang, comb.nov.MycoBank MB833989.
Basionym:Penicillium resinaeZ.T. Qi & H.Z. Kong, Acta Mycol. Sin. 1:
103. 1982.
Notes: Based on an incorrectRPB2sequence,Houbrakenet al.(2014b) considered P. resinae a synonym of Penicillium purpurescens (sect.
Aspergilloides). Penicillium purpurescens predominantly produced simple, monoverticillate conidiophores, whileP.resinaewas originally described to predominantly produce symmetrical biverticillate co-nidiophores.Qi & Kong (1982) classifiedP.resinaein thePenicillium funiculosumseries, nearPenicillium piceum). BothP.funiculosumand P.piceumare currently classified inTalaromycesand it is likely that this species also belongs to this genus. The ex-type strain of P. resinae (CBS 324.83) was re-examined and new sequence data was generated.
Comparison of these sequences show a relationship with Tal. brasi-liensis and Tal. subericola in sect. Trachyspermi (Supplementary Fig. S3).
Talaromyces striatoconidius Houbraken, Frisvad & Samson, nom.
nov.MycoBank MB832566.
Replaced synonym:Phialomyces striatusR.F. Casta~neda & W. Gams, Mycotaxon 42: 239. 1991,non Talaromyces striatus(Raper & Fennell) C.R. Benj. 1955.
Etymology: The epithet refers to the typical striate conidia produced by the species.
Notes: This species was originally described asPhialomyces striatus.
A connection with Penicillium was made and it was suggested that the species takes an intermediate position betweenPhialomycesand Penicillium(Casta~neda & Gams 1991). Molecular data shows that the species belongs to Talaromyces sect. Talaromycesand it is phylo-genetically most closely related toTal.galapagensis,Tal.indigoticus and Tal. rubicundus (Supplementary Fig. S3). Talaromyces striato-conidiuscan be easily differentiated from these related species by its production of striate conidia, a rare feature for most other Talar-omycesspecies. The nameTalaromyces striatus(=Pseudohamigera striata) is already occupied and therefore the nameT.striatoconidius is proposed.
Thermoascus verrucosus(Samson & Tansey) Houbraken, Frisvad &
Samson,comb.nov.MycoBank MB832567.
Basionym: Byssochlamys verrucosa Samson & Tansey, Trans. Brit.
Mycol. Soc. 65: 512. 1975.
Synonym: Paecilomyces verrucosus Samson & Tansey, Trans. Brit.
Mycol. Soc. 65: 512. 1975.
Notes: Samson & Tansey (1975) described B. verrucosa in Byssochlamysbased on phenotypic characters. The conidial state of this species resembles Paecilomyces fulvusand both produce asci from croziers. However, based on an ITS phylogeny,Samsonet al.
(2009) showed that this species is related to Thermoascus. This observation is confirmed in our phylogenetic analysis (Supplementary Fig. S1) and we therefore combine this species in Thermoascus.
Samson & Tansey (1975)already noted that B.verrucosaproduced simple coiled initials, similar to those of Tal. leycettanus and Tal.
purpureus, indicating a position outsideByssochlamys.Thermoascus crustaceus produces, like Th. verrucosus, a paecilomyces-type asexual morph and asci from croziers, and this also indicates a relationship with the genusThermoascus.
Thermoascus yaguchiiHoubraken, Frisvad & Samson,stat.et nom.
nov.MycoBank MB833988.
Replaced synonym:Thermoascus crustaceusvar.verrucosusYaguchi, Someya & Udagawa, Mycoscience 36: 151. 1995,non Thermoascus verrucosus (Samson & Tansey) Houbraken, Frisvad & Samson (this study).
Etymology: In honour of Takashi Yaguchi, a prominent taxonomist studying various genera of Eurotiales and one of the authors that describedThermoascus crustaceusvar.verrucosus.
Synonym: Coonemeria verrucosa (Yaguchi et al.) Mouch., Cryptog.
Mycol. 18: 32. 1997.
Notes:Therrnoascus crustaceusvar.verrucosuswas introduced as a variety because of its verrucose ascospore ornamentation, in contrast to the echinulate ornamentation in Th. crustaceus var. crustaceus.
Sequence data show that this variety represents a distinct species in Thermoascus. Comparison of partialBenAandCaMsequences of CBS 181.67 (a representative strain ofTh.crustaceus) and PF-1160T(= CBS 146343 = IFM 66000), the type ofTh.crustaceusvar.verrucosus, shows a homology of 95.6 and 96.9 %, respectively. The nameThermoascus verrucosusis already occupied and therefore the nameThermoascus yaguchiiis proposed.
Invalid and not accepted species described after 2013
Since the“2014Aspergillus/Penicillium/Talaromyceslists”, around 300 new names have been described inPenicillium,Aspergillusand Talar-omyces. Re-evaluation of these names showed that some of those recently described species were not new to science, incorrectly described or described in the wrong genus. An overview of the not accepted species described after 2013 is given below.
Aspergillus capsici(J.F.H. Beyma) Houbrakenet al., Stud. Mycol. 78:
154. 2014.
Notes: The genusPolypaecilumwas synonymised withAspergillusand thereforePolypaecilum capsiciwas incorrectly combined inAspergillus.
The correct name of Aspergillus capsici (basionym Scopulariopsis capsici J.F.H. Beyma) is Leuconeurospora capsici (J.F.H. Beyma) Mallochet al.
Aspergillus chinensisSamsonet al., Stud. Mycol. 78: 155. 2014.
Notes: The nameAspergillus appendiculatuswas already occupied and therefore the new nameAspergillus chinensiswas introduced for Emer-icella appendiculata(Samsonet al.2014). Phylogenetic and morphologic examination showed thatAspergillus chinensisis identical withAspergillus filifer. We followMatsuzawaet al.(2012), Hubkaet al.(2016a)andChen et al.(2016a)in correcting the nameA.chinensistoAspergillusfilifer Zalaret al.
Aspergillus cicutusP. Singh & P.J. Cotty, Int. J. Food Microbiol. 289:
148. 2018,nom.inval.
Notes: This species is invalidly described: Arts 38.1 (without a description or diagnosis of the taxon), 40.1 (without designation of type specimen), F.5.1 (no identifier cited). None of the markers commonly used inAspergilllustaxonomy (BenA,CaMorRPB2) were available for the representative isolates ofA.cicutus(NRRL 66829, NRRL 66830, NRRL 66831) and only niaD (nitrate reductase) and aflR (aflatoxin pathway transcription factor) sequences were studied (Singh & Cotty 2019). This species could represent a novel species or it is a syno-nym of a sect.Flavimember (e.g.,Aspergillus austwickii).
Aspergillus delacroxiiSamsonet al., Stud. Mycol. 78: 155. 2014.
Notes: From Hubkaet al.(2016a): The nameAspergillus echinulatus was already occupied and therefore the nameAspergillus delacroxiiwas introduced forAspergillus nidulans var. echinulatusFennell & Raper.
The epithet‘‘delacroxii’’was derived from the name of E.G. Delacroix and thus the correct Latin form is“delacroixii”. The spelling‘‘delacroxii’’
could be treated as a correctable orthographical error [Art. 60.1;McNeill et al.(2012)], but the nameA.delacroixiiwas validly proposed twice before, making the nameA.delacroxiiSamsonet al.illegitimate. Even if the epithet‘‘delacroxii’’was grammatically correct, its similarity to the epithet “delacroixii” would likely cause confusion [(Art. 53.3; McNeill et al. (2012)]. We follow Hubkaet al. (2016a)and the correct name forA.nidulansvar.echinulatusisAspergillus spinulosporusHubkaet al.
Aspergillus ferenczii(Varga & Samson) Samsonet al., Stud. Mycol.
78: 155. 2014.
Notes:Samsonet al.(2007a)describedNeosartorya ferencziias a new species in their monograph onAspergillussect.Fumigatiand due to the introduction of the single name nomenclature, the combination in Aspergilluswas subsequently made in 2014 (Samsonet al.2014). In their monograph,Samsonet al.(2007a)were unable to obtain the ex-type cultures ofNeosartorya indohii,N. nishimurae,N.sublevispora,N.
takakii, andN.tsurutae, and these species were therefore treated as doubtful. However, comparison of theBenA,CaMandRPB2sequences of the ex-type strains onA.ferenczii(CBS 121594) andA.sublevisporus (CBS 128796 = IFM 53598) showed that these are similar (BenA, EF669833 vs AB488759: 100 %; CaM, EF669903 vs AB488767:
99.6 %; ITS, EF669977vs MN431376: 100 %;RPB2, EF669764vs MN969095: 99.8 %).Aspergillus ferencziiis regarded to be conspecific withAspergillus sublevisporusSomeyaet al.
Aspergillus frequensHubkaet al., Mycologia 107: 183. 2015.
Notes: The type strains ofA. frequens(NRRL 4578) and A. micro-nesiensis (CBS 138183) have highly similar BenA, CaM andRPB2 sequences. Based on these data, Arzanlouet al. (2016)reducedA.
frequens (Hubka et al. 2015) in synonymy with A. micronesiensis Visagieet al.(Visagieet al.2014a).
Aspergillus korhogoensisA. Carvajal-Campos et al., Toxins 9, 353:
11. 2017.nom.inval.
Notes: An identifier issued by a recognised repository forA. korho-goensiswas not cited in the protologue and this species is therefore invalidly described [Art. 42.1,McNeillet al.(2012)].Frisvadet al.(2019) studied the taxonomy of Aspergillus sect. Flavi and based on their studied specimens, they introduced Aspergillus cerealis. Based on sequence similarity,Aspergillus korhogoensisis treated as a synonym of A.cerealisHoubrakenet al.
Aspergillus latus(Thom & Raper) A.J. Chenet al., Stud. Mycol. 84: 69.
2016.
Notes:Chenet al.(2016a)treatedAspergillus sublatusas a synonym of A.latus. Even thoughA.nidulansvar.latusThom & Raper (1939)is the oldest name of this species, the new combination/status proposed by Chenet al.(2016a)cannot have priority overA.sublatus(oldest name in the species rank) (Hubka et al. 2016a). The correct name for this species isAspergillus sublatusY. Horie.
Aspergillus mangaliensis A. Novakova et al., Mycologia 107: 187.
2015.
Notes: Based on gene concordance and a high sequence homology, Arzanlouet al.(2016)treatedAspergillus mangaliensisas a synonym of A.templicolaVisagieet al.Their conclusion is followed here.
Aspergillus occiafricanusP. Singh & P.J. Cotty, Int. J. Food Microbiol.
289: 148. 2018,nom.inval.
Notes: This species was invalidly described (without an identifier, Art.
F.5.1). NRRL A-11612 was designated as the type. The BenA (MK119746) andCaM(MK119712) sequences derived from the type strain have high homology (99.8 %) with the type ofA.aflatoxiformans (CBS 143679).Aspergillus occiafricanuscan therefore be considered an invalidly described synonym ofA.aflatoxiformansFrisvadet al.
Aspergillus parafelisSuguiet al., J. Clin. Microbiol. 52: 3709. 2014.
Notes:Hubkaet al.(2018a)re-evaluated the species boundaries in the Aspergillus viridinutansspecies complex using a large set of clinical and environmental strains. In their analyses, they showed thatA.parafelis andA.pseudofelisare included in the genetically diverseA.felis line-age. This is in contrary to the results ofSuguiet al.(2014); however, only two isolates of each species were included in their analysis and these isolates did not sufficiently cover the genetic diversity withinA.
felis(Hubkaet al.2018a).
Aspergillus pseudofelisSuguiet al., J. Clin. Microbiol. 52: 3709. 2014.
Notes: This species is a synonym ofA.felisBarrset al.;seeA.parafelis.
Aspergillus similis(Y. Horieet al.) Samsonet al., Stud. Mycol. 78: 157.
2014.
Notes:Samsonet al.(2014)combinedEmericella similisinAspergillus.
Chenet al.(2016a)noted that the ascospore morphology of the ex-type culture (CBS 293.93) is identical to that ofAspergillus violaceus. This close relationship is confirmed by sequence data and we follow the conclusion ofChenet al.(2016a)and treatA.similisas a synonym ofA.
violaceusFennell & Raper.
Aspergillus texensisP. Singhet al., Toxins 10, 513: 6. 2018.
Notes: Molecular phylogenetic analyses ofAspergillus flavus-like iso-lates producing small-sized sclerotia (average sclerotium size < 400μm) collected from across the United States resulted in the discovery of a novel aflatoxin-producing species, named Aspergillus texensis. The phylogenetic position of this species was studied using partialβ-tubulin (0.9 kb), calmodulin (1.2 kb), and nitrate reductase (niaD, 2.1 kb) gene sequences (Singh et al. 2018). The former two gene regions are commonly used in Aspergillus taxonomy; however, the sequenced fragments are longer than those recommended bySamsonet al.(2014).
On the other hand, the nitrate reductase gene is part of the aflatoxin biosynthetic pathway and not commonly used in phylogenetic studies.
Based on the results ofSinghet al.(2018),A.texensisis closely related to A. minisclerotigenes and a previously reported unnamed lineage designated“Lethal Aflatoxicosis Fungus”(LAF). In this study, we re-evaluated the species boundaries of A. minisclerotigenes using a larger set of strains. Single gene phylogenies usingBenA,CaM, and niaD sequences were constructed (using the longer fragments as described bySinghet al.(2018)) (Fig. 20). Thefive representativeA.
texensisstrains clustered together in all analyses, indicating their close genetic relationship. The two included LAF strains (A1168, A1170) causing lethal aflatoxicosis in Kenya (ex maize), clustered together in theBenAandniaDphylogenies, and were closely related in theCaM phylogram. In theBenAphylogram, allA.minisclerotigenes,A.texensis and LAF strains clustered together with high statistical support and there were four minor deviating lineages present. Three main clades are present in theCaM phylogram. One clade contains the A. texensis strains andA.minisclerotigenesstrains DTO 275-E9 and CBS 145094, one clade includes the two LAF strains, CBS 117635Tand four otherA.
minisclerotigenesstrains and one clade with strains only identified asA.
minisclerotigenes. There are also three main clades present in theniaD phylogram; however, the clade containing the LAF strains does not have statistical support. The position ofA.minisclerotigenesCBS 145094 is basal to the other investigated strains. TheA.texensis and the LAF strains clustered together in the combined analysis of the three genes, confirming the results ofSinghet al.(2018). However, with the addition ofA.minisclerotigenesstrains and therefore increasing the genetic di-versity within the analysis, it becomes clear thatA.texensisand the LAF strains fall within the genetic diversity of A. minisclerotigenes. We therefore consider both as synonyms ofA.minisclerotigenesVaamonde et al.
Penicillium attenuatumKirichuk & Pivkin, Mycol. Prog. 16: 21. 2017.
Notes: Kirichuk et al. (2017) introduced Penicillium attenuatum, P. ochotenseandP.piltunenseas new species closely related toP. ant-arcticum(sect.Canescentia, ser.Atroveneta) based on DNA sequences
MT024515 A. minisclerotigenes IBT 32111
BenA
0.001
MK119752 “A. texensis” NRRL 66855 MK119755 “A. texensis” NRRL 66858
MK119753 “A. texensis” NRRL 66856 MK119756 “A. texensis”NRRL 66859 MT024514 A. minisclerotigenes IBT 32094 MK119754 “A. texensis”NRRL 66857
MT024509 A. minisclerotigenes CBS 117634 MT024513 A. minisclerotigenes DTO 045-I9 MK119729 A. minisclerotigenes A-11611 MT024517 A. minisclerotigenes DTO 275-E9 MK119750 Aspergillus sp. A1168 MK119751 Aspergillus sp. A1170
MT024518 A. minisclerotigenes CBS 145094 MK119727 A. minisclerotigenes TAR3N43 MT024510 A. minisclerotigenes FRR 4086 MK119728 A. minisclerotigenes 4-2 MT024516 A. minisclerotigenes CBS 117635T MT024508 A. minisclerotigenes CBS 117633 MT024512 A. minisclerotigenes FRR 5309 MT024511 A. minisclerotigenes FRR 4937 92
MK119743 A. cerealis NRRL 66708 MK119744 A. cerealis NRRL 66709 MK119745 A. cerealis NRRL 66710 89
MK119733 A. flavus AF12 MK119734 A. flavus AF42 MK119735 A. flavus AF70
MK119732 A. flavus AF13 MK119736 A. flavus NRRL 3251 80
MK119739 A. parasiticus NRRL 424 MK119758 A. tamarii NRRL 20818 4x
CaM
0.001
MK119721 “A. texensis”NRRL 66858 MK119719 “A. texensis”NRRL 66856 MK119720 “A. texensis” NRRL 66857 MK119722 “A. texensis”NRRL 66859 MK119718 “A. texensis”NRRL 66855 MT024505 A. minisclerotigenes DTO 275-E9
MT024507 A. minisclerotigenes CBS 145094 MT024503 A. minisclerotigenes IBT 32094 MT024501 A. minisclerotigenes FRR 5309 MT024504 A. minisclerotigenes IBT 32111 MT024500 A. minisclerotigenes FRR 4937 MT024499 A. minisclerotigenes FRR 4086 MT024497 A. minisclerotigenes CBS 117633
MT024498 A. minisclerotigenes CBS 117634 78
MK119693 A. minisclerotigenes TAR3N43 MT024502 A. minisclerotigenes DTO 045-I9 MT024506 A. minisclerotigenes CBS 117635T MK119694 A. minisclerotigenes 4-2 MK119695 A. minisclerotigenes A-11611 MK119716 Aspergillus sp. A1168 MK119717 Aspergillus sp. A1170 MK119701 A. flavus AF70 MK119702 A. flavus NRRL 3251
MK119700 A. flavus AF42 MK119709 A. cerealis NRRL 66708 MK119710 A. cerealis NRRL 66709 MK119711 A. cerealis NRRL 66710
MK119705 A. parasiticus NRRL 424 MK119724 A. tamarii NRRL 20818
4x
Combined
0.001
A. minisclerotigenes CBS 117635T A. minisclerotigenes 4-2
“A. texensis” NRRL 66857
“A. texensis” NRRL 66857